As default, Omega uses the h1.title in region--content.tpl.php.
When using panels, the panel title is used in $title. But what will happen if you want to use your headline inside your panel-layout? A use case might be a project where you have both sidebars in some parts of the project and panels in another part "simulating" a sidebar, for instance with block-styled content in the left panel pane, while main content goes into the right one. Now you will have the headline "within" the main content area on a sidebar display, while it's above the main content area on a panel display. If classic blocks are situated in a sidebar and block-styled content in a sidebar-like panel pane, they will now have a different upper margin relatively to the site header and the headline might also "flip" over the blocks on the left hand side.
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